Noncyclic Photophosphorylation - YouTube

As oxygenic photosynthesis spread through the oceans, everything that could be oxidized by oxygen was, during what is called the (“GOE”), although there may have been multiple dramatic events. The event began as long as three bya and is . The ancient carbon cycle included volcanoes spewing a number of gases into the atmosphere, including hydrogen sulfide, sulfur dioxide, and hydrogen, but carbon dioxide was particularly important. When the continents began forming, carbon dioxide was removed from the atmosphere via water capturing it, , the carbon became combined into calcium carbonate, and plate tectonics subducted the calcium carbonate in the ocean sediments into the crust, which was again released as carbon dioxide in volcanoes.

McGraw-Hill Flash animation illustrating cyclic and non-cyclic photophosphorylation.

Some nonliterate societies do not engage in warfare. They are a vanishingly small proportion of the world’s native societies, but almost without exception, they are not warlike because they are geographically isolated. The most important variable in predicting a society’s level of internal and external violence is male dominance. Monkeys are (matrilineal), and males leave their society of birth to mate, gorillas and chimps are (patrilineal), where females leave their natal society to mate, and humans have both kinds of pre-state societies, along with some . Patrilocal societies are run by gangs of related men, are by far the most violent, engage in the most warfare, and women are subjected to the most violence. Patrilocal societies can also have harems or many “wives” for the alpha males. Patrilocal societies make up nearly 70% of the world’s native cultures that have been documented. Neanderthals and australopiths . The primary determinant of patrilocal or matrilocal residence in humans is the economic contribution of women. In general, where gathering and horticulture brought in more calories than hunting, women had more influence and the society tended to become matrilocal. Those relationships only hold for societies that are not economically centralized. When surplus redistribution appeared, men began to dominate, and the chiefdom was the first step toward state formation. Organized violence only began increasing as states began to form.


Photosynthesis Non Cyclic Photophosphorylation - …

Cyclic DiagramConnections to Electron FlowBoth cyclic and noncyclic photophosphorylation have an electron flow.

Deep-ocean currents, , do not seem to have existed during supercontinental times, and atmospheric oxygen was likely only a few percent at most when the Cryogenian Period began. Canfield’s ocean-oxygenation evidence partly came from testing sulfur isotopes. As with , , and other elements, life prefers the lighter isotope of sulfur, and are two stable isotopes that can be easily tested in sediments. Canfield proposed that in pre-Cryogenian oceanic depths, , which are among Earth’s earliest life forms and produce as its waste product, abounded. Hydrogen sulfide gives rotten eggs their distinctive aroma and is highly toxic to plants and animals, as it . Hydrogen sulfide would react with dissolved iron to form iron pyrite and settle out in the ocean floor, just as the iron oxide did that formed the BIFs. The sulfate-reducing bacteria will enrich the sulfur-32/34 ratio by 3% and did so before the Cryogenian, but the Ediacaran iron pyrite sediments showed a 5% enrichment. A persuasive explanation is recycling sulfur in the oceanic ecosystem, which can only happen in the presence of oxygen.


Non-Cyclic photophosphorylation

During that “,” , , and the rise of grazing and predation had eonic significance. While many critical events in life’s history were unique, one that is not is multicellularity, , and some prokaryotes have multicellular structures, some even with specialized organisms forming colonies. There are , but the primary advantage was size, which would become important in the coming eon of complex life. The rise of complex life might have happened faster than the billion years or so after the basic foundation was set (the complex cell, oxygenic photosynthesis), but geophysical and geochemical processes had their impacts. Perhaps most importantly, the oceans probably did not get oxygenated until just before complex life appeared, as they were sulfidic from 1.8 bya to 700 mya. Atmospheric oxygen is currently thought to have remained at only a few percent at most until about 850 mya, although there are recent arguments that it remained low until only about 420 mya, when large animals began to appear and animals began to colonize land. Just as the atmospheric oxygen content began to rise, then came the biggest ice age in Earth’s history, which probably played a major role in the rise of complex life.

Biology, Eighth Edition (Raven) - Cyclic

About 1 bya, began to decline and microbial photosynthesizers , probably due to predation pressure from , which are eukaryotes. Eating stromatolites may reflect the of , although grazing is really just a form of predation. The difference between grazing and predation is the prey. If the prey is an (it fixes its own carbon, by using energy from either or ), it is called grazing, and if the prey got its carbon from eating autotrophs (such creatures are called ), then it is called . There are other categories of life-form consumption, such as and (eating dead organisms), and there are many instances of . For complex life, the symbiosis between the and its cellular host was the most important one ever.